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Paul Marek

Marek, P.E., T. Tanabe, and S. Sierwald. 2014. A species catalog of the millipede family Xystodesmidae (Diplopoda: Polydesmida), Virginia Museum of Natural History Special Publications, in review.

Kamali, M., P.E. Marek, C. Antonio-Nkondjio, C. Ndo, Z. Tu, F. Simard, and I.V. Sharakhov. 2014. Temporal diversification of major African malaria vectors inferred from multigene phylogenetics, PLosONE, in review.

Marek, P.E., W.A. Shear, and J.E. Bond. 2012. A redescription of the leggiest animal, the millipede Illacme plenipes with notes on its natural history and biogeography (Diplopoda, Siphonophorida, Siphonorhinidae), ZooKeys, 241, 77–112.    [Open access]

Marek, P.E., D.R. Papaj, J. Yeager, S. Molina**, W. Moore. 2011. Bioluminescent aposematism in millipedes, Current Biology, 21, R680-R681.    [Open access]
Video abstract online at Cell Press YouTube channel.

Marek, P.E. 2010. A revision of the Appalachian millipede genus Brachoria Chamberlin, 1939 (Polydesmida, Xystodesmidae, Apheloriini), Zoological Journal of the Linnean Society, 159, 817-889.

Shear, W.A. and P.E. Marek. 2009. Andrognathus hoffmani, n. sp., a second species in the genus and the first species of Andrognathidae from México (Diplopoda, Platydesmida, Andrognathidae). Festschrift in honor of Richard Hoffman, Memoirs of the Virginia Museum of Natural History, 16, 149-158.

Marek, P.E. and J.E. Bond. 2009. A Müllerian mimicry ring in Appalachian millipedes, Proceedings of the National Academy of Sciences, USA. 106, 9755-9760.    [Open access]
Commentary by Merrill, R.M. and C.D. Jiggins. 2009. Müllerian mimicry: sharing the load reduces the legwork, Current Biology, 19, 687-689.    [Open access]

Walker, M.J.**, A.K. Stockman, P.E. Marek, and J.E. Bond. 2009. Pleistocene glacial refugia across the Appalachian Mountains and coastal plain in the millipede genus Narceus: Evidence from population genetic, phylogeographic, and paleoclimatic data, BMC Evolutionary Biology. 9, 25.    [Open access]

Marek, P.E. and J.E. Bond. 2007. A reassessment of apheloriine millipede phylogeny: additional taxa, Bayesian inference, and direct optimization. Zootaxa. 1610, 27-39.

Marek, P.E. and J.E. Bond. 2006. Phylogenetic systematics of the colorful, cyanide-producing millipedes of Appalachia (Polydesmida, Xystodesmidae, Apheloriini) using a total evidence Bayesian approach. Molecular Phylogenetics and Evolution. 41, 704-729.

Marek, P.E. and J.E. Bond. 2006. Biodiversity hotspots: Rediscovery of the world’s leggiest animal. Nature. 441, 707.

Marek, P.E. and D.H. Kavanaugh. 2005. The evolutionary relationships of North American Diplous Motschulsky (Coleoptera: Carabidae: Patrobini) inferred from morphological and molecular evidence. Invertebrate Systematics. 19, 145-168.

Marek, P.E., J.E. Bond, and P. Sierwald. 2003. Rhinocricidae Systematics II: A species catalog of the Rhinocricidae (Diplopoda: Spirobolida) with synonymies. Zootaxa. 308, 1-108.    [Open access]

Bond, J.E. and P.E. Marek. 2003. Rhinocricidae Systematics I: The taxonomic placement of the species of Zipyge Chamberlin, 1925 and Oxypygides Chamberlin, 1922 (Diplopoda: Spirobolida: Rhinocricidae: Oxypyginae). Zootaxa. 292, 1-8.    [Open access]

** student co-authored publications


Non peer-reviewed publications

Marek, P.E. 2013. Xystodesmidae, in: Wikispecies, free species directory, online at

Marek, P.E. 2012. A clever experiment, in: Creatures of Light, American Museum of Natural History iPad app, online at

Marek, P.E. 2012. Environment 4, in: Isopod: The Roly Poly Science Game, Xylem and Phloem LLC iPad app, online at

Marek, P.E. 2010. Brachoria Chamberlin 1939. The Appalachian Mimic Millipedes. Version 16 September 2010. in The Tree of Life Web Project,

Marek, P.E., R.M. Shelley. 2004., An online resource for millipedes, online at